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Check the resistance on the CRE401 that the test button goes to from the JAMMA edge.

To test it, you need to check both the input and output pins between VCC.

For example, if from the JAMMA edge it goes to pin 2, then output is pin 3. Resistance from Pin 1 to pin 2 should be 1k ohm, from pin 1 to pin 3 it should be 1.3k ohm and from pin 2 to pin 3 it should be 300 ohm. If the values are nowhere near that, pull the CRE401 and retest out of circuit. If it's fine out of circuit then it's the custom that handles the i/o but if the CRE401 is bad, much easier fix.
 
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